Mel Brooks got his start in comedy by accidentally flubbing a scene in a play and turning it into a fourth-wall-breaking joke, something that would later become a hallmark of his humor.

Mel Brooks got his first big break in 1950 as a writer on Sid Caesar's 'Your Show of Shows,' a pioneering TV variety show.

Mel Brooks said that learning how to drum as a kid taught him important rules about comedy, including how to structure a joke.

Mel Brooks became an EGOT winner in 2001, meaning he's been awarded an Emmy, a Grammy, an Oscar, and a Tony.

Mel Brooks served in World War II, including a stint clearing landmines at the Battle of the Bulge.

Mel Brooks served as an uncredited producer on David Lynch's film 'The Elephant Man' because he didn't want people to assume it was a lighthearted comedy.

Modern improv comedy, which eschews scripts and emphasizes listening, was developed by University of Chicago students in the early 1950s.

Watch some of the best scenes from Mel Brooks' films, including 'Young Frankenstein,' 'Spaceballs,' 'Robin Hood: Men in Tights,' and more.

Brooks' Broadway adaptation of 'The Producers' was a massive hit when it debuted in 2001, and broke a record for most Tony wins.

Winning a Steve Martin look-alike contest led Bill Nye to pursue comedy

Doing impressions helped Nye realize he enjoyed making people laugh, and he pursued stand-up and comedy writing gigs while working at Boeing after college. On October 3, 1986, he left his engineering job to pursue a full-time career in the entertainment industry.

Mel Brooks wanted Richard Pryor, then a relatively fresh stand-up comic, to star in 1974's 'Blazing Saddles,' but Warner Bros worried that he was not reliable enough for a leading role.

Comedic roasts, an ironic inversion of toasting, took off in the 20th century

Ribbing amongst friends likely goes back to the dawn of man, but formalizing the act caught on with fundraisers held by the Friar’s Club, a New York organization for theater agents. Rather than raising their glasses to good fortune and health, comics eviscerate an “honoree”—along with everyone else on stage—though they usually conclude with an earnest toast.

Will Rogers' political humor was an early predecessor to stand-up

Rogers got his start spinning ropes for circus shows and broke through his association with Florence Zeigfeld's Follies and Frolic. It was his humorous political commentary, though, that would solidify his celebrity status. His goodhearted ribbing of society's "big men" predated the common notion of stand-up comedians as truth-telling jesters.

Before 'SNL,' producer Lorne Michaels was a performer

Michaels was a costar on "The Hart and Lorne Terrific Hour," a popular show on the Canadian network CBC. As these clips show, it was a standard variety show, something Michaels would intentionally react against with "SNL."

Check out a list of some of the best Mel Brooks films, from 'The Producers' and 'History of the World' to 'Spaceballs' and 'Robin Hood: Men in Tights.'

Hear a joke from Charley Case, the first stand-up comedian

Vaudeville performer Case is believed to be the first modern stand-up. While others before him told jokes onstage, Case was the first to do it without props or costuming. This 1909 recording features one of his jokes. The content may be outdated, but the structure can still be found in stand-up sets today.

Mark Twain was an early influence on stand-up

The writer’s live lectures looked a lot like modern stand-up comedy specials. Twain’s world tour in the 1890s featured 90-minute performances in which he stood alone on the stage, delivering long, comedic monologues with his hand perched on his chin.

Lenny Bruce's stand-up challenged US law

Bruce came up in burlesque theaters, but began to find success as stand-up comedy developed a more political and subversive bent. His sets often satirized what he believed to be an oppressively conformist society, which led to his arrest on several occasions.

The first cast members were relatively unknown 'enlightened amateurs'

“SNL” producers aimed to cast relative newcomers for the show’s first season, convinced that amateurs would infuse the show with the countercultural spirit Lorne Michaels envisioned.

Bob Hope's radio shows brought stand-up to the masses

Bob Hope’s 1930s variety shows featured comedic monologues. Unlike the tightly scripted and evergreen sets of vaudeville, Hope’s monologues leaned into topical and political humor, sometimes to the disapproval of his network.

HBO premiered the modern stand-up special in 1975

When they appeared on TV, stand-ups were typically limited to five-minute spots on late-night talk shows, like "The Tonight Show." That changed with the fledgling HBO, which offered comics hourlong slots. The first was "An Evening with Robert Klein," a special that would define "the hour" for decades.

A research lab in Colorado believes comedy is based on ‘benign-violation’

The University of Colorado at Boulder’s Humor Research Lab (which has the decidedly funny abbreviation “HURL”) posits that comedy thrives on relatively harmless violations of dignity or norms.

Conan's 2000 commencement speech offers comfort and humor to young Harvard graduates

In the spring of 2000, famous late-night talk show host and comedian Conan O'Brien gave a hilarious, thoughtful speech to the graduating class at Harvard, where he graduated from in 1985. He warns that once adult life begins, any stupid mistakes made by a Harvard grad will be severely mocked. He also recounts his post-grad failures and how important they were in his success. The speech has been noted ever since as a classic example of both wisdom and foolishness.

George Lucas gave Mel Brooks his blessing to release his 'Star Wars' parody 'Spaceballs,' though he had one condition: 'Spaceballs' could not sell merchandise.

The Borscht Belt was a haven for Jewish vacationers—and comedy fans

New York’s Catskills region resorts were informally named after a dish popular with Ashkenazi Jews. The area offered affordable vacationing, as well as cutting-edge stand-up from future stars like Milton Berle, Mel Brooks, and Joan Rivers.

Stand-up arrived in the late 19th century

Vaudeville theaters featured family-friendly comics, often working in teams, to execute tightly written comedic back-and-forths. Burlesque, vaudeville’s bawdier offshoot, often featured single comics delivering rapid-fire, joke-filled sets.

Steve Martin was vying for love on 'The Dating Game' before he was selling out arenas

In 1968, the 21-year-old Martin was a writer for the Smothers Brothers, who had a popular variety show. He appeared on “The Dating Game” as one of the bachelors, attempting to impress the twin sister of his childhood crush. (Spoiler: It worked.)

One pop-culture historian fooled the nation with a fake origin story of April Fools' Day

In 1983, Joseph Boskin, a professor at Boston University, was interviewed by the Associated Press to discuss the history of April Fools'. Boskin shared a made-up story about a jester who took over Rome for a day and proclaimed that April 1 would henceforth serve as 24 hours of silliness. AP published the fib about "King Kugel," creating buzz and confusion for news outlets around the country.

'The Larry Sanders Show' redefined the American sitcom by removing audience laughter, a move that seemed counterintuitive for a television comedy.

A low-budget, Seattle-based comedy show launched Bill Nye's on-camera career

"Almost Live!" began in 1984 as a Sunday evening talk show before evolving into a sketch comedy show that served as the lead-in for "Saturday Night Live" on NBC's Seattle affiliate station. Besides Nye, TV stars Joel McHale and Lauren Weedman were regular cast members.

Eddie Murphy avoided the show for years thanks to a joke made at his expense

A 1995 segment with David Spade criticized Murphy's unsuccessful film "Vampire in Brooklyn." Muprhy was offended and called the SNL offices to confront Spade. Murphy avoided the show through the 2010s, finally returning in 2015 for the show's 40th anniversary special, where he hugged Spade backstage.

Jim Henson and his Muppets were part of the first season of 'Saturday Night Live' because NBC was hoping to tamp down Lorne Michaels' countercultural plans for the show.

Lenny Bruce was arrested for obscenity seven times

The first was in 1961, after performing at the Jazz Workshop in San Francisco. Subsequent arrests were in Los Angeles, Chicago, and New York. He died of a morphine overdose in 1966, a year and a half after being found guilty of his New York charge.

One actor made a career out of playing Mark Twain

Hal Holbrook toured in the one-man show “Mark Twain Tonight!” for over six decades. In the performances, Holbrook wore Twain’s iconic white suit and offered witticisms similar to Twain’s famous lectures. Holbrook won a Tony Award in 1966 for a Broadway run of the show.

How HBO pioneered the hourlong stand-up comedy special

Stand-up comedy was already on television in the 1970s thanks to Johnny Carson, but comics were limited to clean sets that typically lasted five minutes. HBO, on the other hand, offered stand-ups the opportunity to air their unadulterated material like they would at a club.

An original review of the 1987 parody 'Spaceballs'

On June 24, 1987, Mel Brooks launched "Spaceballs" in theaters—a comedy starring Bill Pullman, John Candy, and Rick Moranis that spoofed the "Star Wars" trilogy and sci-fi tropes in general. The movie has remained a cult classic ever since. Read the Hollywood Reporter's original review of the film here.

Walt Disney’s Laugh-O-Gram Studio featured his earliest innovations in animation

Before he founded his iconic film company, Walt Disney was a young animator in Kansas City, Missouri. After scoring a contract to create animated newsreels for local movie theaters, Disney formed Laugh-O-Gram Studio, where he created innovative animated shorts that set the path for his later work.

Steve Martin and Robin Williams once starred in Samuel Beckett's 'Waiting for Godot,' one of the most well-regarded examples of the existential movement called 'Theater of the Absurd.'

The earliest known play performed in the US was 'The Recruiting Officer'

A London troupe performed a comedy called "The Recruiting Officer" in New York City in 1732.

Michaels doesn’t care when people say ‘SNL’ was funnier when they were younger

"Generally when people talk about the best cast I think, 'Well, that's when they were in high school,'" Michaels told "Today" in 2015. "Because in high school you have the least amount of power you're ever gonna have. ... Staying up with friends later on a Saturday is great, and people attach to a cast."